Department of Agronomy, Zhejiang Key Laboratory of Crop Germplasm, Zhejiang University, Hangzhou, 310058, China. School of Biological Science and Technology, University of Jinan, Jinan, 250022, China. Department of Agronomy, Zhejiang Key Laboratory of Crop Germplasm, Zhejiang University, Hangzhou, 310058, China. zhanggp@zju.edu.cn.